FWS Focus

Overview

Characteristics
Overview

The Lange’s metalmark butterfly is associated with Antioch Dunes, a riverbank dune system that historically reached heights of over 100 feet and may have stretched for miles along the San Joaquin River. Currently, the Lange’s metalmark can only be found on the Antioch Dunes National Wildlife Refuge (ADNWR), the last remnants of the Antioch Dunes. All life stages of Lange’s metalmark butterfly are closely associated with naked-stemmed buckwheat (Eriogonum nudum var. psychicola), which is the primary nectar source for adults, is used to lay eggs and as a larval food plant. The leaves of the larval host plant provide both food and shelter throughout the larval instar phases.

Scientific Name

Apodemia mormo langei
Common Name
Lange's metalmark Butterfly
FWS Category
Insects
Kingdom

Location in Taxonomic Tree

Identification Numbers

TSN:

Characteristics

Characteristic category

Physical Characteristics

Characteristics
Physical Characteristics

Adult butterflies typically begin emerging in early August and may be observed until mid- or late-September. Females lay eggs throughout the adult flight period. This butterfly produces only one brood per year. Females lay eggs singly and in small clusters on larval host plants in late summer. The leaves of the larval host plant provide both food and shelter throughout the larval instar phases. All life stages of Lange’s metalmark are closely tied to a variety of naked-stemmed buckwheat (Eriogonum nudum var. psychicola), which is the primary nectar source for adults, as oviposition sites, and as the larval food plant.

Size & Shape

Lange's metalmark butterfly is a small metalmark butterfly. The wingspan is roughly 25 mm.

Color & Pattern

Lange’s metalmark butterflies have brown and orange coloring on the wings with white markings. The hindwing pattern is characterized by a gray-brown pattern beneath, which can make the butterfly very difficult to see when resting with its wings up in buckwheat foliage. The Lange’s metalmark butterflies is the only metalmark butterfly that occurs at Antioch Dunes, and subtle differences in wing coloration differentiate it from other Mormon metalmark subspecies. Caterpillars are purple and black.

Characteristic category

Life Cycle

Characteristics
Reproduction

Male and female Lange's metalmark butterflies reproduce during the late summer flight period. Males search for females and then mating takes place, usually on plant material. Males and females can stay coupled for a relatively long time (e.g., 2 hours), and are almost motionless during this time. Females lay fertilized eggs within a few days of mating. 

Life Cycle

Adult butterflies typically begin emerging in early August and may be observed until mid- or late September. Females lay eggs throughout the adult flight period. This butterfly produces only one brood per year. Females lay eggs singly and in small clusters on the larval host plant in late summer. The eggs remain dormant until the first part of the rainy season, which usually occurs in the fall, at which time larvae hatch and then crawl to the base of buckwheat plants where they overwinter and feed on the foliage of buckwheat plants. The larvae have been observed feeding at night and have also been observed during daytime. Pupation occurs in mid-summer in the litter at the base of buckwheat plants. The leaves of the larval host plant provide food and shelter throughout the larval instar phases. All life stages of Lange’s metalmark are closely tied to a variety of naked-stemmed buckwheat (Eriogonum nudum var. psychicola), which is the primary nectar source for adults, is used to lay eggs and as a larval food plant.

Life Span

Adult butterflies typically begin emerging in early August and may be observed until mid- or late September. Females lay eggs throughout the adult flight period. This butterfly produces only one brood per year. Females lay eggs singly and in small clusters on the larval host plant in late summer. The eggs remain dormant until the first part of the rainy season, which usually occurs in the fall, at which time larvae hatch and then crawl to the base of buckwheat plants where they overwinter and feed on the foliage of buckwheat plants. The larvae have been observed feeding at night and have also been observed during daytime. Pupation occurs in mid-summer in the litter at the base of buckwheat plants. The leaves of the larval host plant provide food and shelter throughout the larval instar phases. All life stages of the Lange’s metalmark are closely tied to a variety of naked-stemmed buckwheat (Eriogonum nudum var. psychicola), which is the primary nectar source for adults, is used to lay eggs and as a larval food plant.

Characteristic category

Behavior

Characteristics
Behavior

The Lange’s metalmark butterfly (Apodemia mormo langei) is an endangered butterfly associated with the Antioch Dunes ecosystem in Northern California. Currently, the Lange’s metalmark butterfly is only found at Antioch Dunes National Wildlife Refuge, where the last remnants of the Antioch Dunes are preserved. The Lange’s metalmark butterfly is a subspecies of the Mormon metalmark butterfly (Apodemia mormo), which is a member of the Riodinidae family. This is the family that includes the metalmark butterflies. Lange’s metalmark butterflies have brown and orange coloring on the wings with white spotting. The Lange’s metalmark butterfly is the only metalmark butterfly that occurs at Antioch Dunes, and subtle differences in wing coloration differentiate it from other Mormon metalmark butterfly subspecies. Adult butterflies typically begin emerging (eclosing) in early August, and the flight season continues until mid- or late September. The Lange’s metalmark is univoltine, producing only one brood per year. Females lay eggs singly and in small clusters on host plants throughout the flight period. All life stages of Lange’s metalmark butterfly are closely associated with naked-stemmed buckwheat (Eriogonum nudum var. psychicola), which is the primary nectar source for adults, is used to lay eggs and as a larval food plant. The leaves of the larval host plant provide both food and shelter throughout the larval instar phases.

Characteristic category

Similar Species

Characteristics
Similar Species

Lange’s metalmark butterfly is a subspecies of the Mormon metalmark butterfly (Apodemia mormo). All Mormon metalmark butterflies have brown and orange coloring on the wings with white spotting. Lange’s metalmark butterflies is identified by its location as the only Mormon metalmark butterfly subspecies that occurs at Antioch Dunes, along with subtle differences in wing coloration, which differentiate it from other Mormon metalmark butterfly subspecies.

Characteristic category

Habitat

Characteristics
Habitat

All life stages of the Lange’s metalmark butterfly are closely associated with naked-stemmed buckwheat (Eriogonum nudum var. psychicola), which is the primary nectar source for adults, is used to lay eggs and as a larval food plant. The Lange's metalmark butterfly also uses other flowering plants as nectar and food resources, but naked-stemmed buckwheat is considered its primary food plant.  

River or Stream

A natural body of running water.

Urban

Of or relating to cities and the people who live in them.

Characteristic category

Food

Characteristics
Food

Adult butterflies typically begin emerging in early August and may be observed until mid- or late September. Females lay eggs throughout the adult flight period. This butterfly produces only one brood per year. Females lay eggs singly and in small clusters on larval host plants in late summer. The leaves of the larval host plant provide both food and shelter throughout the larval instar phases. All life stages of Lange’s metalmark are closely tied to a variety of naked-stemmed buckwheat (Eriogonum nudum var. psychicola), which is the primary nectar source for adults, is used to lay eggs and as a larval food plant.

Geography

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Timeline

Explore the information available for this taxon's timeline. You can select an event on the timeline to view more information, or cycle through the content available in the carousel below.

8 Items

Listing

Listing

Critical Habitat

Five Year Review

Five Year Review

Five Year Review

Five Year Review

Recovery Plan

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Listing

Oct 14, 1975

Oct 14, 1975 Listing (Endangered)
Proposed Endangered Status for 6 Species of Butterfly in California; 40 FR 48139 48140 (San Bruno el…
  • Publication type: Proposed
Item 1

Listing

Jun 1, 1976

Jun 1, 1976 Listing (Endangered)
Determination That Six Species of Butterflies are Endangered Species; 41 FR 22041 22044 (Lotis blue,…
  • Publication type: Final
Item 2

Critical Habitat

Feb 8, 1977

Feb 8, 1977 Critical Habitat
Proposed Determination of Critical Habitat for Six Butterflies and Two Plants; 42 FR 7972 7976
  • Publication type: Proposed
Item 3

Five Year Review

Feb 14, 2007

Feb 14, 2007 Five Year Review (Information Solicitation)
Initiation of 5-Year Reviews of 58 Species in California and Nevada; Availability of Completed 5-Yea…
  • Publication type: Notice
Item 4

Five Year Review

Mar 25, 2009

Mar 25, 2009 Five Year Review (Completion)
Initiation of 5-Year Reviews of 58 Species in California, Nevada, Arizona, and Utah; Availability of…
  • Publication type: Notice
Item 5

Five Year Review

May 25, 2011

May 25, 2011 Five Year Review (Information Solicitation)
Initiation of 5-Year Reviews of Species in California, Nevada, and the Klamath Basin of Oregon
  • Publication type: Notice
Item 6

Five Year Review

Jun 18, 2018

Jun 18, 2018 Five Year Review (Information Solicitation)
Initiation of 5-Year Status Reviews of 50 Species in California, Nevada, and the Klamath Basin of Or…
  • Publication type: Notice
Item 7

Recovery Plan

Aug 6, 2019

Aug 6, 2019 Recovery Plan (Document Availability (non-FR))
28 Draft Recovery Plan Revisions for 53 Species in the Southeast, Mountain-Prairie, and Pacific Sout…
  • Publication type: Draft
Item 8